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Declan Danaher will celebrate his 250th appearance for London Irish in the Aviva Premiership clash with Harlequins on Sunday.
And while Danaher will reach the personal milestone, at the other end of the scale Anthony Watson will start a Premiership match for the first time.
Both players starred in the 69-26 romp against Mont-de-Marsan in the Amlin Challenge Cup last time out - with director of rugby Brian Smith keeping faith in the same line-up.
There are three changes on the bench, though, with Cai Griffiths, Alex Gray and Darren Allinson coming in.
Harlequins have made three changes for the trip to the Madejski Stadium.
The table-toppers, who recorded back-to-back wins in the Heineken Cup, welcome back Ugo Monye and Tom Casson to the back division, while Tom Guest comes into the back row.
Casson is handed his first start of the season as Jordan Turner-Hall is given the week off to rest an on-going knee problem.
With Nick Evans still sidelined, Ben Botica retains the No 10 jersey.
London Irish: 15 Anthony Watson, 14 Topsy Ojo, 13 Jonathan Joseph, 12 Guy Armitage, 11 Sailosi Tagicakibau, 10 Ian Humphreys, 9 Tomas O'Leary, 8 Chris Hala'ufia 7 Jamie Gibson, 6 Declan Danaher (c), 5 Matt Garvey, 4 Bryn Evans, 3 Halani Aulika, 2 David Paice, 1 Max Lahiff.
Replacements: 16 Scott Lawson, 17 Cai Griffiths, 18 Leo Halavatau, 19 Kieran Low, 20 Ofisa Treviranus, 21 Alex Gray, 22 Steven Shingler, 23 Darren Allinson
Harlequins: 15 Mike Brown, 14 Tom Williams, 13 Matt Hopper, 12 Tom Casson, 11 Ugo Monye, 10 Ben Botica, 9 Danny Care, 1 Joe Marler, 2 Rob Buchanan, 3 James Johnston, 4 Olly Kohn, 5 George Robson, 6 Tom Guest, 7 Chris Robshaw (capt), 8 Nick Easter.
Replacements: 16 Dave Ward, 17 Mark Lambert, 18 Will Collier, 19 Charlie Matthews, 20 Maurie Fa'asavalu, 21 Karl Dickson, 22 Rory Clegg, 23Seb Stegmann
"Sunday will mark a great mile-stone for one of London Irish's finest warriors when Declan Danaher pulls on the London Irish jersey for the 250th time.
"It's an incredible feat of endurance and courage. It will be a tough afternoon for Dec and the boys; we can expect a thorough examination in all departments as Harlequins have a strong pack and some extremely dangerous backline players.
"Whilst we have a healthy respect for our opponents, we have faith in each other and the belief that we can beat any team in this league."
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|Friday 20th December|
|20:00||Sale Sharks vs London Irish|
|Saturday 21st December|
|15:00||Exeter vs Newcastle|
|15:00||Bath vs Harlequins|
|15:00||Wasps vs Northampton|
|15:00||Saracens vs Leicester|
|Friday 27th December|
|TBC||Newcastle vs Wasps|
|Saturday 28th December|
|15:00||Northampton vs Bath|
|15:00||Worcester vs Saracens|
|15:00||Harlequins vs Exeter|
|15:00||Leicester vs Sale Sharks|
|Friday 3rd January|
|19:45||Newcastle vs Sale Sharks|
|Saturday 4th January|
|15:00||London Irish vs Worcester|
|15:00||Gloucester vs Saracens|
|15:00||Leicester vs Bath|
|Sunday 5th January|
|15:00||Wasps vs Exeter|
|Friday 7th February|
|19:45||Worcester vs Leicester|
|20:00||Sale Sharks vs Gloucester|
|Saturday 8th February|
|15:00||Harlequins vs Wasps|
|15:00||Bath vs Newcastle|
|15:00||Exeter vs Northampton|
|Sunday 9th February|
|15:00||Saracens vs London Irish|
|Friday 14th February|
|20:00||Sale Sharks vs Saracens|
|Saturday 15th February|
|15:00||Wasps vs London Irish|
|15:00||Northampton vs Worcester|
|15:00||Leicester vs Gloucester|
|Sunday 1st December|
|Newcastle 9 - 35 Harlequins|
|Saturday 30th November|
|Bath 21 - 16 Exeter|
|Worcester 10 - 33 Northampton|
|London Irish 12 - 19 Wasps|
|Saracens 24 - 19 Sale Sharks|
|Friday 29th November|
|Gloucester 17 - 22 Leicester|
|Sunday 24th November|
|Wasps 5 - 28 Bath|
|Saturday 23rd November|
|Northampton 18 - 0 Newcastle|
|Harlequins 27 - 19 Gloucester|
|Exeter 9 - 16 Saracens|
|Leicester 20 - 11 London Irish|
|Friday 22nd November|
|Sale Sharks 26 - 10 Worcester|
|Sunday 3rd November|
|Saracens 40 - 3 Newcastle|
|London Irish 14 - 19 Northampton|
|Saturday 2nd November|
|Leicester 16 - 23 Harlequins|
|Gloucester 30 - 32 Wasps|
|Friday 1st November|
|Sale Sharks 16 - 18 Exeter|
|Worcester 6 - 21 Bath|
|Sunday 27th October|
|Newcastle 13 - 11 London Irish|
|Wasps 22 - 12 Leicester|
|Saturday 26th October|
|Northampton 41 - 20 Saracens|
|Exeter 40 - 6 Worcester|
|Harlequins 24 - 3 Sale Sharks|
|Friday 25th October|
|Bath 15 - 13 Gloucester|
Read the thoughts and opinions of Stuart Barnes with skysports.com