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Declan Danaher will celebrate his 250th appearance for London Irish in the Aviva Premiership clash with Harlequins on Sunday.
And while Danaher will reach the personal milestone, at the other end of the scale Anthony Watson will start a Premiership match for the first time.
Both players starred in the 69-26 romp against Mont-de-Marsan in the Amlin Challenge Cup last time out - with director of rugby Brian Smith keeping faith in the same line-up.
There are three changes on the bench, though, with Cai Griffiths, Alex Gray and Darren Allinson coming in.
Harlequins have made three changes for the trip to the Madejski Stadium.
The table-toppers, who recorded back-to-back wins in the Heineken Cup, welcome back Ugo Monye and Tom Casson to the back division, while Tom Guest comes into the back row.
Casson is handed his first start of the season as Jordan Turner-Hall is given the week off to rest an on-going knee problem.
With Nick Evans still sidelined, Ben Botica retains the No 10 jersey.
London Irish: 15 Anthony Watson, 14 Topsy Ojo, 13 Jonathan Joseph, 12 Guy Armitage, 11 Sailosi Tagicakibau, 10 Ian Humphreys, 9 Tomas O'Leary, 8 Chris Hala'ufia 7 Jamie Gibson, 6 Declan Danaher (c), 5 Matt Garvey, 4 Bryn Evans, 3 Halani Aulika, 2 David Paice, 1 Max Lahiff.
Replacements: 16 Scott Lawson, 17 Cai Griffiths, 18 Leo Halavatau, 19 Kieran Low, 20 Ofisa Treviranus, 21 Alex Gray, 22 Steven Shingler, 23 Darren Allinson
Harlequins: 15 Mike Brown, 14 Tom Williams, 13 Matt Hopper, 12 Tom Casson, 11 Ugo Monye, 10 Ben Botica, 9 Danny Care, 1 Joe Marler, 2 Rob Buchanan, 3 James Johnston, 4 Olly Kohn, 5 George Robson, 6 Tom Guest, 7 Chris Robshaw (capt), 8 Nick Easter.
Replacements: 16 Dave Ward, 17 Mark Lambert, 18 Will Collier, 19 Charlie Matthews, 20 Maurie Fa'asavalu, 21 Karl Dickson, 22 Rory Clegg, 23Seb Stegmann
"Sunday will mark a great mile-stone for one of London Irish's finest warriors when Declan Danaher pulls on the London Irish jersey for the 250th time.
"It's an incredible feat of endurance and courage. It will be a tough afternoon for Dec and the boys; we can expect a thorough examination in all departments as Harlequins have a strong pack and some extremely dangerous backline players.
"Whilst we have a healthy respect for our opponents, we have faith in each other and the belief that we can beat any team in this league."
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|Friday 21st March|
|19:45||Worcester vs Wasps|
|Saturday 22nd March|
|14:00||Sale Sharks vs Northampton|
|15:00||Gloucester vs Newcastle|
|15:15||Saracens vs Harlequins|
|17:30||London Irish vs Bath|
|Sunday 23rd March|
|14:00||Leicester vs Exeter|
|Friday 28th March|
|19:45||Bath vs Sale|
|Saturday 29th March|
|15:00||Wasps vs Saracens|
|15:00||Harlequins vs London Irish|
|15:00||Exeter vs Gloucester|
|15:15||Northampton vs Leicester|
|Sunday 30th March|
|14:00||Newcastle vs Worcester|
|Friday 11th April|
|19:45||Sale vs Harlequins|
|Saturday 12th April|
|15:00||Worcester vs Exeter|
|15:00||Leicester vs Wasps|
|15:15||Gloucester vs Bath|
|Sunday 13th April|
|14:00||Saracens vs Northampton|
|15:00||London Irish vs Newcastle|
|Friday 18th April|
|19:45||Harlequins vs Leicester|
|Saturday 19th April|
|15:00||Bath vs Worcester|
|15:00||Exeter vs Sale Sharks|
|15:15||Wasps vs Gloucester|
|Sunday 20th April|
|14:00||Northampton vs London Irish|
|15:00||Newcastle vs Saracens|
|Saturday 3rd May|
|15:00||Bath vs Northampton|
|Sunday 2nd March|
|Newcastle 18 - 41 Leicester|
|Saturday 1st March|
|Northampton 39 - 13 Gloucester|
|Exeter 18 - 0 London Irish|
|Harlequins 21 - 20 Worcester|
|Wasps 17 - 21 Sale Sharks|
|Friday 28th February|
|Bath 10 - 23 Saracens|
|Sunday 23rd February|
|Saracens 23 - 10 Exeter|
|Newcastle 16 - 22 Northampton|
|London Irish 15 - 20 Leicester|
|Saturday 22nd February|
|Worcester 12 - 24 Sale Sharks|
|Bath 32 - 25 Wasps|
|Gloucester 25 - 20 Harlequins|
|Sunday 16th February|
|Leicester 11 - 8 Gloucester|
|Saturday 15th February|
|Exeter 23 - 27 Bath|
|Wasps 20 - 23 London Irish|
|Northampton 30 - 14 Worcester|
|Harlequins 18 - 14 Newcastle|
|Friday 14th February|
|Sale Sharks 10 - 15 Saracens|
|Sunday 9th February|
|Harlequins 11 - 10 Wasps|
|Saturday 8th February|
|Exeter 16 - 17 Northampton|
|Saracens 13 - 22 London Irish|
|Bath 24 - 6 Newcastle|
|Friday 7th February|
|Sale Sharks 24 - 19 Gloucester|
|Worcester 22 - 23 Leicester|
Read the thoughts and opinions of Stuart Barnes with skysports.com