Leicester sign Dominic Ryan, Sione Kalamafoni, Joe Ford and Jonah Holmes

Leicester have announced the signings of Dominic Ryan, Sione Kalamafoni, Joe Ford and Jonah Holmes.

Irish flanker Ryan is joining from Leinster, while Tonga back row Kalamafoni makes the switch from the Tigers' Premiership rivals Gloucester.

Fly-half Ford and wing Holmes, the leading points-scorer and try-scorer respectively in last year's Championship, both arrive from Leeds.

Leicester head coach, Matt O'Connor, worked with Irish Wolfhounds cap Ryan at Leinster and said: "Dom is an experienced professional who plays all across the back-row.

"He wins collisions on both sides of the ball, and he comes from an environment that understands the narrative of winning.

"Joe is an experienced fly-half whose game-management is exceptional. He brings others into the game and in all his previous clubs has made the team tick at a high level.

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"Jonah is a talented young player who knows where the try-line is, as his record shows. He has huge growth left in his game and we know he will get that development at Tigers."

Of Kalamafoni, he said: "Sione is a strong and powerful ball-carrier from the back-row. That physical presence on both sides of the ball is something we have been looking to add to our squad."

Ford will team up with his brother George at Welford Road next season.